Preheat the oven to 350°F. Spray a 9x13-inch baking pan with nonstick spray.
In a bowl, using an electric hand mixer or in the bowl of a stand mixer, beat together the 15.25 ounce box lemon cake mix, 1 cup whole milk, ½ cup (1 stick) unsalted butter, melted and cooled, and 3 large eggs for 2 minutes on medium-high speed. Pour the batter into your prepared pan.
Bake on the middle rack for about 20 to 25 minutes, or until a toothpick inserted in the center comes out clean. Let the cake cool for 30 minutes.
Filling:
Using the back of a wooden spoon or a wide straw, poke holes all over the cake.
Whisk together the 2 cups whole milk and 3.4 ounces INSTANT lemon pudding mix, then quickly pour the mixture over the cake, making sure to fill the holes.
Refrigerate the cake for 1 hour to set.
Topping:
Spread the 8 ounce tub whipped topping, thawed over the cake, then sprinkle the lemon zest from two lemons on top.
Notes
Please refer to my FAQ’s (Frequently Asked Questions) and ingredient list above for other substitutions or for the answers to the most common questions.